“Our results suggest that mark–recapture estimates may have been negatively biased due to limited spatial sampling. We observed large numbers of bears summering in southeastern WH, an area not regularly sampled by mark–recapture.” Stapleton et al. 2014.
We’ve seen the results of this 2011 study before, in government report format. But now it’s been revamped, peer-reviewed and published in a respected scientific journal – it actually came out in February, without fanfare, but I’ve only just come across it.
Some excerpts below, with conclusions that should raise some eyebrows.